fungus – Should lenses be stored without their caps inside a dry cabinet?

From what I have learn, there a potential for lenses to get fungus when stored for a long time in a dark and humid area.

For this reason, it is recommended to store them in a dry cabinet that has some way to control humidity between 35% and 45%, and be translucent so that non direct sunlight can pass through.

I have seem lots of people store their lenses with the caps on, to prevent scratches when they bump into each other inside the box, but on the other side, this prevents the ambient light to penetrate the lenses.

So, what you think, should the lenses be stored in a dry cabinate with or without their caps?